Extrachromosomal DNA associates with nuclear condensates and reorganizes chromatin structures to enhance oncogenic transcription.

Document Type

Article

Publication Date

12-8-2025

Keywords

JGM, SS1, Humans, Chromatin, Transcription, Genetic, DNA, Circular, Neoplasms, Cell Line, Tumor, Gene Expression Regulation, Neoplastic, Cell Nucleus, Epigenesis, Genetic, Carcinogenesis, Animals, Oncogenes, Mice

JAX Source

Cancer Cell. 2025;43(12):2191–205 e6.

ISSN

1878-3686

PMID

40972571

DOI

https://doi.org/10.1016/j.ccell.2025.08.008

Abstract

Extrachromosomal, circular DNA (ecDNA) is a prevalent oncogenic alteration in cancer genomes, often associated with aggressive tumor behavior and poor patient outcome. While previous studies proposed a chromatin-based mobile enhancer model for ecDNA-driven oncogenesis, its precise mechanism and impact remains unclear across diverse cancer types. Our study, utilizing advanced multi-omics profiling, epigenetic editing, and imaging approaches in three cancer models, reveals that ecDNA hubs are an integrated part of nuclear condensates and exhibit cancer-type specific chromatin connectivity. Epigenetic silencing of the ecDNA-specific regulatory modules or chemically disrupting nuclear condensates breaks down ecDNA hubs, displaces MED1 co-activator binding, inhibits oncogenic transcription, and promotes cell death. These findings substantiate the trans-activator function of ecDNA and underscore a structural mechanism driving oncogenesis. This refined understanding expands our views of oncogene regulation and opens potential avenues for alternative therapeutic strategies in cancer treatment.

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